Honey Bee A Tribute to Randolf Menzel
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For other uses, see Honey bee disambiguation. Main article: Apis mellifera. Main article: Africanized bee. Main articles: Pollination management and List of crop plants pollinated by bees.
Main article: Beekeeping. Main article: Colony collapse disorder. Main article: Honey. Main article: Beeswax.
Main article: Bee pollen. Main article: Propolis. Main article: Royal jelly. Main article: Drone bee. Main article: Worker bee.
Main article: Queen bee. Main article: Worker policing. Main article: Bee sting. This section needs additional citations for verification. Please help improve this article by adding citations to reliable sources.
Unsourced material may be challenged and removed. Main article: Bee learning and communication. Main article: Bee mythology.
Giant honey bee A. Play media. Apis mellifera on rock rose Cistus in Oakland, California. Online Etymology Dictionary, Douglas Harper. Retrieved Snodgrass Anatomy of the Honey Bee.
Cornell University Press. Retrieved February 26, Entomological Society of America. Retrieved February 21, Tree of Life Web Project.
Palmer Archived from the original PDF on February 29, Engel; I. Hinojosa-Diaz; A. Rasnitsyn Proceedings of the California Academy of Sciences.
Retrieved 9 July Engel Journal of Hymenoptera Research. Encyclopedia of Life. Arias; Walter S. Sheppard Molecular Phylogenetics and Evolution.
Maria C. Bee World. Gloag; Denis L. Anderson; Benjamin P. Oldroyd Systematic Entomology. Randall Journal of the Kansas Entomological Society. June 26, Archived from the original on Engel, Ismael A.
Gould; Carol Grant Gould The Honey Bee. Scientific American Library. Archived from the original on 4 March Retrieved 12 March In some places the researchers considered, wild insects were pollinating most of the plants despite rented honey bees being present.
Honeybees also spread exotic plants and transmit pathogens, both of which have been shown to harm other pollinators. Elite Daily.
Quote checkers like this one, and this one agree. But debunking its message? That's more complicated. For another, the statement is hyperbolic and wrong and Einstein was rarely wrong.
Business Insider. Grasses and grassland ecology. Oxford: Oxford University Press. Behavioral Ecology and Sociobiology.
Colony losses have occurred periodically throughout history. Fungus, mites, and starvation have all been thought to be the cause of the deaths. The first recorded collapses were called "May Disease" in Colorado in and Journal of Invertebrate Pathology.
The Independent. Retrieved 2 July Archived from the original on 6 February Retrieved 23 October Bee Culture. Molecular Ecology.
Retrieved 14 February Retrieved 15 May Home technologies and practices for small agricultural producers, UN Food and Agriculture Organization. National Geographic.
National Library of Medicine. January 19, BMC Dermatol. Z Bienenforsch. ID ; "fish protein" ID ; acidic water-based, non-alcoholic flavoured beverages containing calcium in the range of 0.
EFSA Journal. Food and Drug Administration. Apr 5, Visual Dictionary, QA International. Retrieved 18 May PerfectBee LLC. Beesource Beekeeping.
Ecology and Evolution. H Insectes Sociaux. Journal of Evolutionary Biology. Kirk Bibcode : Natur.
Bibcode : PNAS.. Behavior Ecology and Sociobiology. Insectes Soc. An Introduction to Behavioral Ecology.
West Sussex: Wiley-Blackwell, Davidson College. Archived from the original on October 2, Retrieved June 1, Bibcode : NW BBC News. Retrieved July 5, Randall; Engel, Michael S.
Honeybees of Asia. Neuroscience Bulletin. W; Bone, R. The queen's job is simple—laying the eggs that will spawn the hive's next generation of bees.
There is usually only a single queen in a hive. Queens also regulate the hive's activities by producing chemicals that guide the behavior of the other bees.
Male bees are called drones—the third class of honeybee. Several hundred drones live in each hive during the spring and summer, but they are expelled for the winter months when the hive goes into a lean survival mode.
Bees live on stored honey and pollen all winter, and cluster into a ball to conserve warmth.
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Details vary among the different species of honey bees, but common features include:. The worker bees rotate through the cluster from the outside to the inside so that no bee gets too cold.
The colder the weather is outside, the more compact the cluster becomes. During winter, they consume their stored honey to produce body heat. During the summer, however, this is achieved through fanning and water evaporation from water collected in various fields.
Of all the honey bee species, only A. Species of Apis are generalist floral visitors, and pollinate many species of flowering plants, but because of their "generalized" nature, they do so inefficiently.
Without specialized adaptations for specific flowers, their ability to reach pollen and nectar is often limited. What's more, their tendency to visit all species in a given area means that the pollen they carry for any one species is often very diluted.
As such, they can provide some pollination to many plants, especially non-native crops, but most native plants have some native pollinator that is far more effective at pollinating that species.
Western honey bees are often described as being essential to all human food production, leading to claims that without their pollination, all of humanity would starve, or even die out.
In fact, many important crops need no insect pollination at all. Soybeans are self-pollinated. Rice , wheat , sorghum , and maize are all wind pollinated , as are most other grasses.
Similarly, no crops originating in the New World depend on the western honey bee Apis mellifera at all, as the insect is invasive, having been brought over with colonists in the last few centuries.
Tomatoes , peppers , squash , and all other New World crops evolved with native pollinators like squash bees , bumble bees , and other native bees.
The stingless bees mentioned by Jefferson are distant relatives of the honey bees, in the genus Melipona.
Honey bees obtain all of their nutritional requirements from a diverse combination of pollen and nectar. Pollen is the only natural protein source for honey bees.
Adult worker honey bees consume 3. Of these amino acids, honey bees require highest concentrations of leucine, isoleucine, and valine, however elevated concentrations of arginine and lysine are required for brood rearing.
Pyridoxine is the most prevalent B vitamin found in royal jelly and concentrations vary throughout the foraging season with lowest concentrations found in May and highest concentrations found in July and August.
Honey bees lacking dietary pyridoxine were unable to rear brood. Pollen is also a lipid source for honey bees ranging from 0.
Fat-soluble vitamins A, D, E, and K are not considered essential but have shown to significantly improve the number of brood reared. Nurse bees have the ability to selectively transfer sterols to larvae through brood food.
Nectar is collected by foraging worker bees as a source of water and carbohydrates in the form of sucrose.
The dominant monosaccharides in honey bee diets are fructose and glucose but the most common circulating sugar in hemolymph is trehalose which is a disaccharide consisting of two glucose molecules.
Honey bees require water to maintain osmotic homeostasis, prepare liquid brood food, and to cool the hive through evaporation.
A colony's water needs can generally be met by nectar foraging as it has high water content. Occasionally on hot days or when nectar is limited, foragers will collect water from streams or ponds to meet the needs of the hive.
The only domesticated species of honey bee are A. In Japan, where mellifera is vulnerable to local hornets and disease, the Japanese honey bee a. Modern hives also enable beekeepers to transport bees, moving from field to field as the crop needs pollinating and allowing the beekeeper to charge for the pollination services they provide, revising the historical role of the self-employed beekeeper, and favoring large-scale commercial operations.
Bees of various types other than honey bees are also domesticated and used for pollination or other means around the world, including Tetragonula iridipennis in India, the blue orchard bee for tree nut and fruit pollination in the United States, and a number of species of Bombus bumblebees for pollination in various regions globally, such as tomatoes , which are not effectively pollinated by honey bees.
Primarily in places where western honey bees were imported by humans, periodic collapses in western honey bee populations have occurred at least since the late 19th century.
This has been dubbed "colony collapse disorder" CCD and was at first unexplained. Larval stages of the moth G. Eggs are laid within the hive, and the larvae that hatch tunnel through and destroy the honeycombs that contain bee larva and their honey stores.
The tunnels they create are lined with silk, which entangles and starves emerging bees. Destruction of honeycombs also result in honey leaking and being wasted.
Both G. To manage the mite, temperature treatments are possible, but also distorts wax of the honeycombs. Chemical fumigants, particularly CO 2 , are also used.
Honey is the complex substance made when bees ingest nectar, process it, and store the substance into honey combs.
Worker bees of a certain age secrete beeswax from a series of exocrine glands on their abdomens. As with honey, beeswax is gathered by humans for various purposes such as candle making, waterproofing, soap and cosmetics manufacturing, pharmaceuticals, art, furniture polish and more.
Bees collect pollen in their pollen baskets and carry it back to the hive. Worker bees combine pollen, honey and glandular secretions and allow it to ferment in the comb to make bee bread.
The fermentation process releases additional nutrients from the pollen and can produce antibiotics and fatty acids which inhibit spoilage.
In the hive, pollen is used as a protein source necessary during brood-rearing. In certain environments, excess pollen can be collected from the hives of A.
The product is used as a health supplement. It has been used with moderate success as a source of pollen for hand pollination.
Bee brood — the eggs, larvae or pupae of honey bees — is nutritious and seen as a delicacy in countries such as Indonesia,  Mexico, Thailand, and many African countries; it has been consumed since ancient times by the Chinese and Egyptians.
Propolis is a resinous mixture collected by honey bees from tree buds, sap flows or other botanical sources, which is used as a sealant for unwanted open spaces in the hive.
Royal jelly is a honey bee secretion used to nourish the larvae. Honey bees have three castes : drones , workers, and queens.
Drones are typically haploid , having only one set of chromosomes , and primarily exist for the purpose of reproduction. There are rare instances of diploid drone larvae.
This phenomenon usually arises when there is more than two generations of brother-sister mating. In developing bees, if the conditions are that the individual is heterozygous for the csd gene, they will develop into females.
If the conditions are so that the individual is hemizygous or homozygous for the csd gene, they will develop into males. The instances where the individual is homozygous at this gene are the instances of diploid males.
They do not defend the hive or kill intruders, and do not have a stinger. Workers have two sets of chromosomes. Workers typically develop in 21 days.
A typical colony may contain as many as 60, worker bees. Their duties change upon the age of the bee in the following order beginning with cleaning out their own cell after eating through their capped brood cell : feed brood, receive nectar, clean hive, guard duty, and foraging.
Workers have morphological specializations, including the pollen basket corbicula ,  abdominal glands that produce beeswax, brood-feeding glands, and barbs on the sting.
Under certain conditions for example, if the colony becomes queenless , a worker may develop ovaries. Worker honey bees perform different behavioural tasks that cause them to be exposed to different local environments.
Queen honey bees are created when worker bees feed a single female larvae an exclusive diet of a food called " royal jelly ".
In addition to the greater size of the queen, she has a functional set of ovaries, and a spermatheca, which stores and maintains sperm after she has mated.
Apis queens practice polyandry , with one female mating with multiple males. The highest documented mating frequency for an Apis queen is in Apis nigrocincta , where queens mate with an extremely high number of males with observed numbers of different matings ranging from 42 to 69 drones per queen.
Once mated, queens may lay up to 2, eggs per day. When a fertile female worker produces drones, a conflict arises between her interests and those of the queen.
The worker shares half her genes with the drone and one-quarter with her brothers, favouring her offspring over those of the queen.
The queen shares half her genes with her sons and one-quarter with the sons of fertile female workers. This relationship leads to a phenomenon known as "worker policing".
In these rare situations, other worker bees in the hive who are genetically more related to the queen's sons than those of the fertile workers will patrol the hive and remove worker-laid eggs.
Another form of worker-based policing is aggression toward fertile females. In very rare instances workers subvert the policing mechanisms of the hive, laying eggs which are removed at a lower rate by other workers; this is known as anarchic syndrome.
Anarchic workers can activate their ovaries at a higher rate and contribute a greater proportion of males to the hive.
Although an increase in the number of drones would decrease the overall productivity of the hive, the reproductive fitness of the drones' mother would increase.
Anarchic syndrome is an example of selection working in opposite directions at the individual and group levels for the stability of the hive.
Under ordinary circumstances the death or removal of a queen increases reproduction in workers, and a significant proportion of workers will have active ovaries in the absence of a queen.
The workers of the hive produce a last batch of drones before the hive eventually collapses. Although during this period worker policing is usually absent, in certain groups of bees it continues.
According to the strategy of kin selection , worker policing is not favored if a queen does not mate multiple times. Workers would be related by three-quarters of their genes, and the difference in relationship between sons of the queen and those of the other workers would decrease.
The benefit of policing is negated, and policing is less favored. Experiments confirming this hypothesis have shown a correlation between higher mating rates and increased rates of worker policing in many species of social hymenoptera.
All honey bees live in colonies where the workers sting intruders as a form of defense, and alarmed bees release a pheromone that stimulates the attack response in other bees.
The different species of honey bees are distinguished from all other bee species and virtually all other Hymenoptera by the possession of small barbs on the sting, but these barbs are found only in the worker bees.
The sting apparatus, including the barbs, may have evolved specifically in response to predation by vertebrates, as the barbs do not usually function and the sting apparatus does not detach unless the sting is embedded in fleshy tissue.
While the sting can also penetrate the membranes between joints in the exoskeleton of other insects and is used in fights between queens , in the case of Apis cerana japonica , defense against larger insects such as predatory wasps e.
Asian giant hornet is usually performed by surrounding the intruder with a mass of defending worker bees, which vibrate their muscles vigorously to raise the temperature of the intruder to a lethal level "balling".
Defense can vary based on the habitat of the bee. In the case of those honey bee species with open combs e.
Another act of defense against nest invaders, particularly wasps, is "body shaking," a violent and pendulum like swaying of the abdomen, performed by worker bees.
The stings of honey bees are barbed and therefore embed themselves into the sting site, and the sting apparatus has its own musculature and ganglion which keep delivering venom even after detachment.
The embedded stinger continues to emit additional alarm pheromone after it has torn loose; other defensive workers are thereby attracted to the sting site.
The worker dies after the sting becomes lodged and is subsequently torn loose from the bee's abdomen. The honey bee's venom, known as apitoxin , carries several active components, the most abundant of which is melittin ,  and the most biologically active are enzymes , particularly phospholipase A2.
Honey bee venom is under laboratory and clinical research for its potential properties and uses in reducing risks for adverse events from bee venom therapy ,  rheumatoid arthritis ,  and use as an immunotherapy for protection against allergies from insect stings.
With an increased number of honey bees in a specific area due to beekeeping, Western honey bees as an invasive species and native wild bees often have to compete for the limited habitat and food sources available,  and Western honey bees may become defensive in response to the seasonal arrival of competition from other colonies, particularly Africanized bees which may be on the offence and defence year round due to their tropical origin.
Honey bees are known to communicate through many different chemicals and odors, as is common in insects. They also rely on a sophisticated dance language that conveys information about the distance and direction to a specific location typically a nutritional source, e.
The dance language is also used during the process of reproductive fission, or swarming, when scouts communicate the location and quality of nesting sites.
The details of the signalling being used vary from species to species; for example, the two smallest species, Apis andreniformis and A.
Carniolan honey bees Apis mellifera carnica use their antennae asymmetrically for social interactions, with a strong lateral preference to use their right antennae.
There has been speculation as to honey bee consciousness. Honey bees have a tiny structure that appears similar to a human midbrain, so if it functions the same way they may possibly be able to achieve a small amount of simple awareness of their bodies.
The bee was used as a symbol of government by Emperor Napoleon I of France. The priestess at Delphi was the "Delphic Bee". The Quran has a Sura chapter titled " The Bee ".
It is named after honey bees, and contains a comparison of the industry and adaptability of honey bees to the industry of man. And your Lord taught the honey bee to build its cells in hills, on trees, and in men's habitations; Then to eat of all the produce of the earth , and find with skill the spacious paths of its Lord: there issues from within their bodies a drink of varying colours, wherein is healing for men: verily in this is a Sign for those who give thought.
Workers are the only bees that most people ever see. These bees are females that are not sexually developed. Workers forage for food pollen and nectar from flowers , build and protect the hive, clean, circulate air by beating their wings, and perform many other societal functions.
The queen's job is simple—laying the eggs that will spawn the hive's next generation of bees. There is usually only a single queen in a hive. Queens also regulate the hive's activities by producing chemicals that guide the behavior of the other bees.
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Lemon Drop Sequin Mix. Amethyst Crystal Sequin Mix. Vitamin Sea Sequin Mix. It's all in the little details We are a small family-owned business based in Oregon.Although an increase in the number of drones would decrease the overall productivity of the hive, the reproductive fitness of Www Casinoclub De drones' mother would increase. Archived from the Vandal PDF on February 29, Scientific Reports. The worker shares half her genes with the drone and one-quarter with her brothers, favouring her offspring over those of the queen. We are a small family-owned business based in Oregon. H; Kim, J. Encarsia formosa Ladybird Parasitoid wasp. Utah State Library. Visual Dictionary, QA International. Die fleißigen Mitarbeiter honey Bee™ Fahrradanhänger für Kinder ist die perfekte Balance zwischen Wert und Vielseitigkeit, ohne Kompromisse bei der. Abstract. During cocoon construction the predominant movement made by larvae of the three honey bee castes is a forward somersault which is made dorsal side. Vitellogenin in Honey Bee Behavior and Lifespan Foraging Honey Bees: How Foragers Determine and Transmit Information About Feeding Site Locations. Within a honey bee (Apis mellifera) colony there are individuals who In recognition of a corpse within the nest, worker bees probably use chemical cues. Burley Fahrradanhänger Honey Bee - Children Trailercycle, red, x x cm, jet-rent.be: Bekleidung.
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